Skip to main content


Dr Paul Rose

Dr Paul Rose


 Not Known

 Washington Singer 226


Washington Singer Laboratories, University of Exeter, Perry Road, Prince of Wales Road, Exeter, EX4 4QG, UK


I am a lecturer in Psychology, based in the Centre for Research in Animal Behaviour. My research predominantly centres on captive wild animals in zoos, aquariums and other ex situ facilities. I use evidence from behavioural studies to inform animal husbandry and management, and to improve welfare.

I have written the book "The behavioural biology of zoo animals" and I am currently working on another volume specifically relating to zoo animal welfare. 

I welcome approaches from PhD students on prospective topics and I am happy to discuss specific research questions, appropriate to a PhD with interested students. 

Visit my eprofile pages for more information about my research interests.

Research group links

Back to top


Research interests

My main research interests focus on:

  • Assessment of animal welfare using behavioural data
  • Evidence for appropriate animal management husbandry
  • Social network approaches and assessment of animal welfare 
  • Measurement of behaviour using remote technology 
  • Defining the relevance of natural behaviour to zoo animal welfare
  • Conservation of ex situ populations 

I am also interested in the role of nature connectivity to human wellbeing and the importance of blue and green experiences (i.e. being in nature) to human quality of life. I also research the messaging that conservation organisations broadcast on their social media platforms. 

I am open to discussing research proposals on animal behaviour and animal welfare pertaining to captive wild animals or other populations of conservation importance. If you are interested in contacting me about a possible research topic, please send me an email with an outline of your ideas. 

Back to top



Rose P (2022). The behavioural biology of zoo animals. Abstract.

Journal articles

Rice T, Reed A, Badman-King A, Hurn S, Rose P (In Press). A Desirable Privation: exploring silence as a mode of zoo visiting. TRACE: Journal for Human-Animal Studies Abstract.
Croft DP, Rose PE (In Press). Evidence of directed interactions between individuals in captive flamingo flocks. Wildfowl, 65, 121-132.
Rice T, Badman-King A, Reed A, Hurn S, Rose P (In Press). Listening After the Animals: sound and pastoral care in the zoo. Journal of the Royal Anthropological Institute
Rice T, Reed A, Badman-King A, Hurn S, Rose P (In Press). Listening to the zoo: challenging zoo visiting conventions. Ethnos: journal of anthropology
Rose P, Brereton JE, Croft DP (In Press). Measuring welfare in captive flamingos: activity patterns and exhibit usage in zoo-housed birds. Applied Animal Behaviour Science
Rose P, Lloyd I, Brereton JE, Croft D (In Press). Patterns of nocturnal activity in captive greater flamingos. Zoo Biology
Rose P, Croft DP (In Press). Quantifying the social structure of a large captive flock of greater flamingos (Phoenicopterus roseus): potential implications for management in captivity. Behavioural Processes
Rose P, Croft DP (In Press). Social bonds in a flock bird. Species differences and seasonality in social structure in captive flamingo flocks over a 12-month period. Applied Animal Behaviour Science
Rose P, Croft DP (In Press). The potential of social network analysis as a tool for the management of zoo animals. Animal Welfare
Loader A, Rose P (2023). Age-Related Change in the Association Choices of Two Species of Juvenile Flamingos. Animals, 13(16), 2623-2623. Abstract.
Brereton J, Rose P (2023). An evaluation of the role of ‘biological evidence’ in zoo and aquarium enrichment practices. Animal Welfare, 31(1), 13-26. Abstract.
Kelly R, Rose PE (2023). Assessing the impact of environmental enrichment on behavior in understudied armadillo species: a case study. Zoo Biology, 43(1), 100-109. Abstract.
Mooney A, McCall K, Bastow S, Rose P (2023). Changes in Environment and Management Practices Improve Foot Health in Zoo-Housed Flamingos. Animals, 13(15), 2483-2483. Abstract.
Rose P, Riley L (2023). Five ways to wellbeing at the zoo: improving human health and connection to nature. Frontiers in Psychology, 14 Abstract.
Mooney A, Teare JA, Staerk J, Smeele SQ, Rose P, Edell RH, King CE, Conrad L, Buckley YM (2023). Flock size and structure influence reproductive success in four species of flamingo in 540 captive populations worldwide. Zoo Biology, 42(3), 343-356. Abstract.
Rose P (2023). Identifying Essential Elements of Good Giraffe Welfare—Can We Use Knowledge of a Species’ Fundamental Needs to Develop Welfare-Focussed Husbandry?. Journal of Zoological and Botanical Gardens, 4(3), 549-566. Abstract.
McCully FR, Rose PE (2023). Individual personality predicts social network assemblages in a colonial bird. Scientific Reports, 13(1). Abstract.
Claydon M, Brereton J, Rose P (2023). Never be mute about bird welfare: Swanning around with environmental enrichment. Zoo Biology, 43(1), 83-91. Abstract.
Greenwell PJ, Riley LM, Lemos de Figueiredo R, Brereton JE, Mooney A, Rose PE (2023). The Societal Value of the Modern Zoo: a Commentary on How Zoos can Positively Impact on Human Populations Locally and Globally. Journal of Zoological and Botanical Gardens, 4(1), 53-69. Abstract.
Brereton JE, Rose PE (2023). The Zone Overlap Index: a new measure of shared resource use in the zoo. Zoo Biology, 42(6), 811-817.
Des Pallieres CG, Rose PE (2023). Two’s company, three species is a crowd? a webcam-based study of the behavioural effects of mixed-species groupings in the wild and in the zoo. PLOS ONE, 18(4), e0284221-e0284221. Abstract.
George AJ, Rose PE (2023). Wing condition does not negatively impact time budget, enclosure usage, or social bonds in a flock of both full-winged and flight-restrained greater flamingos. Zoo Biol, 42(6), 766-779. Abstract.  Author URL.
Wood KA, Lacey R, Rose PE (2022). Assessing trade-offs in avian behaviour using remotely collected data from a webcam. PLOS ONE, 17(7), e0271257-e0271257. Abstract.
McConnell H, Brereton J, Rice T, Rose P (2022). Do Birds of a Feather Always Flock Together? Assessing Differences in Group and Individual Zoo Enclosure Usage by Comparing Commonly Available Methods. Journal of Zoological and Botanical Gardens, 3(1), 71-88. Abstract.
Rose P, Reed A, Hurn S, Badman-King A, Rice T (2022). Does the sound environment influence the behaviour of zoo-housed birds? a preliminary investigation of ten species across two zoos. Behavioural Processes, 203, 104763-104763.
Munday C, Rose P (2022). Environmental and Social Influences on the Behaviour of Free-Living Mandarin Ducks in Richmond Park. Animals, 12(19), 2554-2554. Abstract.
Rose P, Roper A, Banks S, Giorgio C, Timms M, Vaughan P, Hatch S, Halpin S, Thomas J, O’Brien M, et al (2022). Evaluation of the time-activity budgets of captive ducks (Anatidae) compared to wild counterparts. Applied Animal Behaviour Science, 251, 105626-105626.
Rose PE, Riley LM (2022). Expanding the role of the future zoo: Wellbeing should become the fifth aim for modern zoos. Frontiers in Psychology, 13 Abstract.
Kidd P, Ford S, Rose PE (2022). Exploring the Effect of the COVID-19 Zoo Closure Period on Flamingo Behaviour and Enclosure Use at Two Institutions. Birds, 3(1), 117-137. Abstract.
Knoch S, Whiteside MA, Madden J, Rose P, Fawcett T (2022). Hot-headed peckers: thermographic changes during aggression among juvenile pheasants (Phasianus colchicus). Philosophical Transactions of the Royal Society B: Biological Sciences, 377, 20200442-20200442.
Rose P (2022). Observation of an Attempted Forced Copulation within a Captive Flock of Greater Flamingos (Phoenicopterus roseus). Birds, 3(4), 374-382. Abstract.
Rose PE, Chapman J, Brereton JE, Riley LM (2022). What’s Black and White and Pink all Over? Lesser Flamingo Nocturnal Behaviour Captured by Remote Cameras. Journal of Zoological and Botanical Gardens, 3(4), 624-640. Abstract.
Harley JJ, O’Hara L, Rose PE (2021). A Global Survey of Current Zoo Housing and Husbandry Practices for Fossa: a Preliminary Review. Journal of Zoological and Botanical Gardens, 2(3), 388-405. Abstract.
Boultwood J, O’Brien M, Rose P (2021). Bold Frogs or Shy Toads? How Did the COVID-19 Closure of Zoological Organisations Affect Amphibian Activity?. Animals, 11(7), 1982-1982. Abstract.
Rose PE, Riley LM (2021). Conducting Behavioural Research in the Zoo: a Guide to Ten Important Methods, Concepts and Theories. Journal of Zoological and Botanical Gardens, 2(3), 421-444. Abstract.
Llewellyn T, Rose PE (2021). Education is Entertainment? Zoo Science Communication on YouTube. Journal of Zoological and Botanical Gardens, 2(2), 250-264. Abstract.
Rose P (2021). Evidence for Aviculture: Identifying Research Needs to Advance the Role of Ex Situ Bird Populations in Conservation Initiatives and Collection Planning. Birds, 2, 77-95.
Kidd P, Rose P (2021). Influences of Rearing Environment on Behaviour and Welfare of Captive Chilean Flamingos: a Case Study on Foster-Reared and Parent-Reared Birds. Journal of Zoological and Botanical Gardens, 2(2), 174-206. Abstract.
Lewton J, Rose PE (2021). Social networks research in ex situ populations: Patterns, trends, and future directions for conservation‐focused behavioral research. Zoo Biology, 40(6), 493-502. Abstract.
Grace MK, Akçakaya HR, Bennett EL, Brooks TM, Heath A, Hedges S, Hilton-Taylor C, Hoffmann M, Hochkirch A, Jenkins R, et al (2021). Testing a global standard for quantifying species recovery and assessing conservation impact. Conserv Biol, 35(6), 1833-1849. Abstract.  Author URL.
Brereton JE, Fryer J, Rose PE (2021). Understanding sociality and behavior change associated with a nesting event in a captive flock of great white pelicans. Zoo Biology, 40(5), 386-397. Abstract.
Rose P, Badman‐King A, Hurn S, Rice T (2021). Visitor presence and a changing soundscape, alongside environmental parameters, can predict enclosure usage in captive flamingos. Zoo Biology, 40(5), 363-375. Abstract.
Wood KA, Ham P, Scales J, Wyeth E, Rose PE (2020). Aggressive behavioural interactions between swans (<i>Cygnus</i>spp.) and other waterbirds during winter: a webcam-based study. AVIAN RESEARCH, 11(1).  Author URL.
Riley LM, Rose PE (2020). Concepts, applications, uses and evaluation of environmental enrichment: Perceptions of zoo professionals. JOURNAL OF ZOO AND AQUARIUM RESEARCH, 8(1), 18-28.  Author URL.
Rose PE, Croft DP (2020). Evaluating the social networks of four flocks of captive flamingos over a five-year period: Temporal, environmental, group and health influences on assortment. Behavioural Processes, 175 Abstract.
Lewton J, Rose PE (2020). Evaluating the social structure of captive Rothschild’s giraffes (Giraffa camelopardalis rothschildi): Relevance to animal management and animal welfare. Journal of Applied Animal Welfare Science, 23(2), 178-192. Abstract.
Rose P, Rowden L (2020). Specialised for the Swamp, Catered for in Captivity? a Cross-Institutional Evaluation of Captive Husbandry for Two Species of Lechwe. Animals, 10, 1874-1874. Abstract.
Lang GPSA, Rose PE, Nash SM, Riley LM (2020). The nocturnal activity of a commonly housed rodent: How African pygmy dormice (Graphiurus murinus) respond to an enriched environment?. Journal of Veterinary Behavior, 38, 82-88.
Rose P, O’Brien M (2020). Welfare Assessment for Captive Anseriformes: a Guide for Practitioners and Animal Keepers. Animals, 10(7), 1132-1132. Abstract.
Rose P, Soole L (2020). What influences aggression and foraging activity in social birds? Measuring individual, group and environmental characteristics. Ethology Abstract.
Rose PE, Scales JS, Brereton JE (2020). Why the “Visitor Effect” is Complicated. Unraveling Individual Animal, Visitor Number, and Climatic Influences on Behavior, Space Use and Interactions with Keepers—A Case Study on Captive Hornbills. Frontiers in Veterinary Science, 7
Rose P (2019). Evaluating the behaviour of Andean Flamingos Phoenicoparrus andinus and James’s Flamingos P. jamesi in captivity: comparing species and flocks using multiple methods. Wildfowl, 69, 70-92.
Rose P (2019). The use of Qualitative Behavioural Assessment to zoo welfare measurement and animal husbandry change. Journal of Zoo and Aquarium Research, 7, 150-161.
Rose P (2019). What’s new from the zoo? an analysis of ten years of zoo-themed research output. Palgrave Communications, 5
Rose P, Hunt K, Riley L (2018). Animals in an online world; an evaluation of how zoological collections use social media. Journal of Zoo & Aquarium Research, 6, 57-62.
Regaiolli B, Sandri C, Rose PE, Vallarin V, Spiezio C (2018). Investigating parental care behaviour in same-sex pairing of zoo greater flamingo (<i>Phoenicopterus roseus</i>). PeerJ, 6, e5227-e5227. Abstract.
Hulbert AJ, Hunt KA, Rose PE (2017). A multi-zoo investigation of nutrient provision for captive red-crested turacos. Zoo Biology, 36(2), 152-160. Abstract.
Rose PE, Nash SM, Riley LM (2017). Moving forward with zoo welfare assessment: a response to Cooke (2017). Journal of Veterinary Behavior: Clinical Applications and Research, 22, 75-77. Abstract.
Rose P, Nash SM, Riley LM (2017). To pace or not to pace? a review of what Abnormal Repetitive Behavior tells us about zoo animal management. Journal of Veterinary Behavior: Clinical Applications and Research, 20, 11-11.
Wood KA, Ponting J, D'Costa N, Newth JL, Rose PE, Glazov P, Rees EC (2017). Understanding intrinsic and extrinsic drivers of aggressive behaviour in waterbird assemblages: a meta-analysis. Animal Behaviour, 126, 209-216. Abstract.
Rose P, Rowden L (2016). A global survey of banteng (Bos javanicus) housing and husbandry. Zoo Biology
Rose P (2016). Developing flamingo husbandry practices through workshop communication. Journal of Zoo & Aquarium Research, 4, 115-121.
Rose PE, Croft DP, Lee R (2014). A review of captive flamingo (Phoenicopteridae) welfare: a synthesis of current knowledge and future directions. International Zoo Yearbook, 48(1), 139-155. Abstract.
Rose PE, Croft DP, Lee R (2014). A review of captive flamingo (Phoenicopteridae) welfare: a synthesis of current knowledge and future directions. International Zoo Yearbook
Rose PE, Roffe SM (2013). A Case Study of Malayan Tapir (Tapirus indicus) Husbandry Practice Across 10 Zoological Collections. Zoo Biology, 32(3), 347-356. Abstract.


Rose P, Huffman BA (2022). Behavioural biology and the future zoo: Overall conclusions. In  (Ed) The Behavioural Biology of Zoo Animals, 331-341.
Riley LM, Díez-León M, Rose P (2022). Behavioural biology and zoo animal welfare: for the future. In  (Ed) The Behavioural Biology of Zoo Animals, 285-292.
Rose P (2022). Introduction to the behavioural biology of the zoo. In  (Ed) The Behavioural Biology of Zoo Animals, 3-10.
Stevens C, Fiddes M, Rose P (2022). The behavioural biology of freshwater fishes. In  (Ed) The Behavioural Biology of Zoo Animals, 237-252.
Hickey I, Rose P, Rowden L (2022). The behavioural biology of ungulates and elephants. In  (Ed) The Behavioural Biology of Zoo Animals, 67-82.
Rose P, Mooney A, Klass J (2022). The behavioural biology of waterbirds. In  (Ed) The Behavioural Biology of Zoo Animals, 153-170.
Rose P (2022). Threatened Flamingos: Conservation to Keep the Firebird from Being Extinguished. In  (Ed) Imperiled: the Encyclopedia of Conservation, Elsevier, 29-49.
Rose PE (2020). Flamingo Social Behavior and Flock Dynamics. In  (Ed) Encyclopedia of Avian Science Volume 1-4, 903-959. Abstract.
Rose P (2018). Ensuring a Good Quality of Life in the Zoo: Underpinning Welfare-Positive Animal Management with Ecological Evidence. In Berger M, Corbett S (Eds.) Zoo Animals: Husbandry, Welfare and Public Interactions, New York, USA: Nova Science Publishers. Abstract.
Rose PE (2016). Flamingo social behavior and flock dynamics. In  (Ed) Flamingos: Behavior, Biology, and Relationship with Humans, 123-180. Abstract.

Back to top

External Engagement and Impact

Advice to Government, Parliament, devolved and English regional administrations, other national, international, regional or local agencies; advice to non-government organisations and to private practice

I currently sit on Defra's Zoos Expert Committee. 

External Examiner Positions

I am an Exeternal Examiner for the University of Chester and for Salford University.

Back to top


My main teaching specialisms are in animal behaviour (including behavioural ecology and applied animal behaviour science), as well as research methods and approaches to quantitative studies. 

In the 2023-2024 academic year, I am teaching:

PSY1105 Introduction to Animal Behaviour & Evolution

PSYM225 Biological Psychology

PSYM222 Research Methods & Conceptual Issues in Psychology

PSYM205 Advances and Methods in Animal Behaviour



Back to top

Edit Profile